Jennifer Trump boards a TGV train in Paris to chase a story about BioGenesis. Ken Carpenter, the conductor, starts the trip to Bordeaux. A sick woman gets on, bites people, and soon the train is full of infected monsters. Jennifer and Ken team up with Chloe, a runaway teen, Marie, a pregnant woman, and others to stay alive.
They fight through cars, find out BioGenesis made the virus, and try to stop the train before it crashes. Ken and others die to save them. In Bordeaux, the station falls to the infected, but Jennifer, Chloe, and Marie escape with proof of the truth. They reach safety, ready to tell the world, but the danger isn't over.
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